菌落小且厚實(shí)，圓整，光滑、平旦；菌體桿狀，大小1.0μm×2.5–4.0μm，革蘭氏陰性。參考文獻(xiàn)：Lee, E. and Manoil, C. 1994. Mutations eliminating the protein export function of a membrane-spanning sequence. J. Biol. Chem. 269: 28822–28828. 基因型：(ara-leu)7697 _lacX74 _phoA20 galE galK thi rpsE rpoB argE(am) recA1
